Google Scholar Profile

See my Google Scholar profile.


Since 2002, my students and I have published ~200 total scientific papers and book chapters (~186 indexed on ISI Web of Science WOS) that have been cited 9,404 times on WOS and ~15,694 times on Google Scholar. My h-index on WOS is 48 and Google Scholar 59 (June 2021).

(Underlined names are student and post-doctoral fellow publications, ** are undergraduate thesis papers, ++ denotes publication awards)

In Review, Revision, & Press

  1. Hebblewhite, M., Hilty, J.A., Locke, H., Chester, C., Johns, D., Kehm, G., Francis, W.L., 2021. Yellowstone to Yukon Conservation Initiative: Large-Landscape Conservation from Theory to Results in Twenty Years. Frontiers in Ecology and Environment, In Review.
  2. Paterson, T. Proffitt, K., DeCesare, N.J., Hebblewhite, M., Gude, J. 2021. Evaluating the summer landscapes of predation risk and forage quality for elk (Cervus canadensis). Oikos, In review.
  3. Forzely, M., Rotella, J., Proffitt, K., Garrott, R., Jimenez, B., Eacker, D.R., Hebblewhite, M. 2021. Effects of Increasing Mountain Lion Harvest on Annual Elk Calf Survival, Ecosphere, In Review.
  4. Whittington, J., R. Baron, M. Hebblewhite, A. T. Ford, and J. Paczkowski. 2021. Towns and trails drive carnivore connectivity using a step selection approach. Movement Ecology, In Review.
  5. Williams, S., M. Hebblewhite, H. Martin, J. Killeen, J. Berg, K. M. MacAulay, P. Smolko, and E. H. Merrill. 2021. Predation Risk Drives Long-Term Shifts in Migratory Behavior and Demography in a Large Herbivore Population. Journal of Animal Ecology, In Review.
  6. MacAulay, K. M., E. Spilker, J. Berg, M. Hebblewhite, and E. H. Merrill. 2021. Predicting multi-predator risk to elk (Cervus canadensis) using scats: Are migrant elk exposed to different predation risk? Ecology & Evolution, In Review.
  7. Palm, E., M. Suitor, K. Joly, J. Herriges, A. Kelly, D. Hervieux, K. Russell, T. Bentzen, N. Larter, and M. Hebblewhite. 2021. Increasing fire frequency and severity will increase habitat loss for a boreal forest indicator species. Ecological Applications, In Revision.
  8. Martin, H., M. Hebblewhite, and E. H. Merrill. 2021. Ideal free migration? Testing for density-dependent migration in partially migratory elk. Ecology, In Revision.
  9. Hessami, M., Martin, H., Whittington, J., Hebblewhite, M., Steenweg, R., Flowers, M., Merrill, E.H. Estimating Neonatal Elk Calf Survival with Remote Cameras using Royle-Nichol’s Abundance Models. Wildlife Society Bulletin, In Revision.
  10. Lamb, C., R. Willson, C. Richter, N. Owens-Beek, J. Napoleon, B. Muir, S. McNay, E. Lavis, M. Hebblewhite, L. Giguere, T. Dokkie, B. S., and A. Ford. 2021. Indigenous-led conservation: pathways to recovery for the nearly extirpated Klinse-Za mountain caribou. Ecological Applications, In Revision.
  11. McNay, R. S., C. Lamb, L. Giguere, S. Williams, H. Martin, M. Hebblewhite, and G. D. Sutherland. 2021. Demographic responses of nearly extirpated endangered mountain caribou to recovery actions in central British Columbia. Ecological Applications, In Revision.



  1.  Normandeau, J., C. Cassady-St.Clair, S. Kutz, M. Hebblewhite, and E. H. Merrill. 2021. What makes elk tick: winter tick grooming behavior in wild elk (Cervus canadensis). Journal of Mammalogy, In Press.
  2. Labadie, G., McLoughlin, P., Hebblewhite, M., Fortin, D. 2021. Insect-mediated apparent competition between mammals in a boreal food web. Proceedings of the National Academy of Sciences, In Press.
  3. Berg, J. E., J. Reimer, P. Smolko, H. Bohm, M. Hebblewhite, and Merrill, E.H. 2021. Mother’s movements: shifts in calving area selection by Partially migratory elk. Journal of Wildlife Management, In Press.
  4. Kauffman, M. J., F. Cagnacci, S. Chamaillé-Jammes, Hebblewhite, J. G. C. Hopcraft, J. A. Merkle, T. Mueller, A. Mysterud, W. Peters, C. Roettger, A. Steingisser, J. E. Meacham, K. Abera, J. Adamczewski, E. O. Aikens, H. Bartlam-Brooks, E. Bennit, J. Berger, C. Boyd, S. D. Côté, L. Debeffe, A. S. Dekrout, N. Dejid, E. Donadio, L. Dziba, W. F. Fagan, C. Fischer, S. Focardi, J. M. Fryxell, R. W. S. Fynn, C. Geremia, B. A. González, A. Gunn, E. Gurarie, M. Heurich, J. A. Hilty, M. A. Hurley, A. Johnson, K. Joly, P. Kaczensky, C. J. Kendall, P. Kochkarev, L. Kolpaschikov, R. Kowalczyk, F. van Langevelde, V. Binbin, A. L. Lobora, A. Loison, T. H. Madiri, D. Mallon, P. Marchand, R. A. Medellin, E. Meisingset, M. E.H., A. D. Middleton, K. L. Monteith, M. D. Morjan, T. M. Morrison, S. Mumme, R. Naidoo, A. J. Novaro, O. J.O., K. A. Olson, Y. Oteng-Yeboah, R. J. A. Ovejero, N. Owen-Smith, A. Paasivaara, C. Packer, D. Panchenko, L. Pedrotti, A. J. Plumptre, C. M. Rolandsen, S. Said, A. Salemgareyev, A. P. Savchenko, P. Savchenko, H. Sawyer, M. Selebatso, M. Skroch, E. J. Solberg, J. A. Stabach, O. Strand, M. J. Suitor, Y. Tachiki, A. M. Trainor, A. Tshipa, M. Z. Virani, C. Vynne, S. Ward, G. Wittemyer, W. Xu, and S. Zuther. 2021. Mapping out a future for ungulate migrations: Limited mapping of migrations hampers conservation. Science 372:566 - 569.
  5. Suraci, J. P., K. M. Gaynor, M. L. Allen, P. Alexander, J. S. Brashares, S. Cendejas-Zarelli, K. Crooks, L. M. Elbroch, T. Forrester, A. M. Green, J. Haight, N. C. Harris, Hebblewhite, F. Isbell, B. Johnston, R. Kays, P. E. Lendrum, J. S. Lewis, A. McInturff, W. McShea, T. W. Murphy, M. S. Palmer, A. Parsons, M. A. Parsons, M. E. Pendergast, C. Pekins, L. R. Prugh, K. A. Sager-Fradkin, S. Schuttler, C. H. Sekercioglu, B. Shepherd, L. Whipple, J. Whittington, G. Wittemyer, and C. C. Wilmers. 2021. Disturbance type and species life history predict mammal responses to humans. Global Change Biology, Early View,


  1. Jakes, A., P. F. Jones, C. Gates, K. Kunkel, S. Olimb, and Hebblewhite. 2020. Multi-scale assessment of migration habitat for pronghorn. PLoS One 15:e0241042.
  2. **Heinle, K., Eby, L., Muhlfeld, C.C., Steed, A., Jones, L., D’Angelo, V., Whiteley, A.R., Hebblewhite, M. 2020. Influence of water temperature and biotic interactions on the distribution of Westslope Cutthroat Trout (Oncorhynchus clarkii lewisi) under climate change. Canadian Journal of Fisheries and Aquatic Sciences, 78: 444-456.
  3. Moeller, A., J. Nowak, L. Neufeld, M. Bradley, M. Manseau, P. Wilson, S. McFarlane, P. M. Lukacs, and Hebblewhite. 2020. Integrating counts, telemetry, and non-invasive DNA data to improve demographic monitoring of an endangered species. Ecosphere, 12(5): e03443.
  4. Davidson, S. C., G. Bohrer, E. Gurarie, J. Jennewein, S. LaPoint, P. J. Mahoney, N. T. Boelman, J. Eitel, L. R. Prugh, L. Vierling, L. Grier, A. J. H. Meddens, R. Y. Oliver, R. Kays, R. Wikelski, T. Aarvak, J. T. Ackerman, J. A. Alves, E. Bayne, B. Bedrosian, J. L. Belant, A. B. Berdahl, A. B. Berlin, D. Berteaux, J. Bêty, D. Boiko, T. L. Booms, B. L. Borg, S. Boutin, W. S. Boyd, K. Brides, S. Brown, K. K. Burnham, D. Cabot, M. Casazza, K. Christie, E. H. Craig, e. al, and Hebblewhite. 2020. Ecological insights from three decades of animal movement tracking across a changing Arctic. Science 370:712-715.
  5. Nagy‐Reis, M., M. Dickie, A. M. Calvert, Hebblewhite, D. Hervieux, D. R. Seip, S. L. Gilbert, O. Venter, C. DeMars, S. Boutin, and R. Serrouya. 2021. Habitat loss accelerates for the endangered woodland caribou in western Canada. Conservation Science and Practice.
  6. Williams, S. H., R. Steenweg, T. Hegel, M. Russell, D. Hervieux, and Hebblewhite. 2021. Habitat loss on seasonal migratory range imperils an endangered ungulate. Ecological Solutions and Evidence 2. 1- 12.
  7. Mahoney, P., K. Joly, B. L. Borg, M. S. Sorum, T. Rinaldi, G. Howard, D. Saalfeld, D. Lathan, K. Allicia, B. Mangapine, C. Lambert, L. Neufeld, Hebblewhite, and N. T. Boelman. 2020. Denning phenology and reproductive success of wolves in response to climate signals. Global Change Biology 15:125001.
  8. Jennewein, J. S., Hebblewhite, P. Mahoney, S. Gilbert, A. J. H. Meddens, N. T. Boelman, K. Joly, K. Jones, K. A. Kellie, S. Brainerd, L. A. Vierling, and J. U. H. Eitel. 2020. Behavioral modifications by a large-northern herbivore to mitigate warming conditions. Movevment Ecology 8:39.
  9. Hebblewhite, M., and J. Whittington. 2020. Wolves Without Borders: Transboundary Survival of Wolves in Banff National Park over Three Decades. Global Ecology and Conservation 24:e01293.
  10. Clark, T. J., S. Horne, M. Hebblewhite, and A. D. Luis. 2020. Stochastic predation exposes prey to predator pits and local extinction. Oikos 130:300-309.
  11. Clark, T. J., and Hebblewhite. 2020. Predator control may not increase ungulate populations in the future: A formal meta‐analysis. Journal of Applied Ecology 58:812-824.
  12. Normandeau, J., H. Martin, E. H. Merrill, and Hebblewhite. 2021. Potential case of pseudo-hermaphroditism in Elk (Cervus canadensis) in Alberta, Canada. Canadian Field Naturalist 134:241-247.
  13. Proffitt, K., R. Garrott, J. A. Gude, Hebblewhite, B. Jimenez, J. T. Paterson, and J. Rotella. 2020. Integrated carnivore-ungulate management: a case study in west-central Montana. Wildlife Monographs, In Press. Wildlife Monographs 206:1–28.
  14. Pigeon, K. E., G. Stenhouse, Hebblewhite, M. Musiani, L. Neufeld, J. Cranston, D. MacEarney, and F. Schmiegelow. 2020. The density of anthropogenic features explains seasonal and behavior-based functional responses in selection of linear features by a social predator. Scientific Reports, 11: 11437.
  15. Aikens, E.O., Mysterud, A., Merkle, J., Cagnacci, F., Rivrud, I.M., Hebblewhite, M., Hurley, M.A., Peters, W., Bergen, S., De Groeve, J., Dwinnel, S.P.H., Gehr, B., Heurich, M., Hewison, M.A., Jarnemo, A., Kjellander, P., Kroschel, M., Licoppe, A., Linnell, J., Merrill, E.H., Middleton, A.D., Morellet, N., Neufeld, L., Ortega, A.C., Parker, K.L., Pedrotti, L., Proffitt, K.M., Said, S., Sawyer, H., Scurlock, B.M., Singer, J., Stent, P., Sustr, P., Szkorupa, T., Monteith, K.L., Kauffman, M.J. 2020. Wave-like patterns of plant phenology determine ungulate migration tactics across northern temperate ecosystems, Current Biology, 30: 1- 6. org/10.1016/j.cub.2020.06.032
  16. Merrill, E.H., Killeen, J., Pettit, J., Trottier, M., Martin, H., Berg, J., Bohm, H., Eggeman, S., Hebblewhite, M. Density-dependent foraging behaviors on sympatric winter ranges in a partially migratory elk (Cervus canadensis) population. Frontiers in Ecology and Evolution, 8: 269. doi: 10.3389/fevo.2020.00269
  17. **Metz, M., SunderRaj, J., Smith, D., Stahler, D., Kohl, M., Cassidy, K., Hebblewhite, M. 2020. Accounting for imperfect detection in observational studies: modeling wolf sightability in Yellowstone National Park. Ecosphere, 11(6): 031452 1002/ecs2.3152. += Undergraduate thesis publication.
  18. Jones, P.F., Jakes, A.F., Eacker, D.R., Hebblewhite, M. 2020. Annual pronghorn survival of a partially migratory population. Journal of Wildlife Management, 84: 1114-1126,
  19. Normandeau, J., Kutz, S.J., Hebblewhite, M., Merrill, E.H. 2020. Living with liver flukes: does migration matter? IJP: Parasites and Wildlife, 12: 76- 84.
  20. Hurley, M., Hebblewhite, M., Gaillard, J.-M. 2020. Competition for Safe Real Estate Rather than Food Drives Density-Dependent Juvenile Survival in a Large Herbivore. Ecology and Evolution, 00: 1-12.
  21. Palm, E., Aerin, Jacob, Fluker, S., Nesbitt, H., and Hebblewhite, M. Beyond designation: the long road to protecting critical habitat for species at risk. Conservation Science and Practice, 2: e219.
  22. Geremia, C., Merkle, J.A., Eacker, D.R., Wallen, R.L., White, P.J., Hebblewhite, M., Kauffman, M.J. 2020. Reply to Joseph M. Craine: Bison redefine what it means to move to find food. Proceedings of the National Academy of Sciences, 117: 9171 – 9172
  23. Fortin, D., Hebblewhite, M., McLoughlin, P.D., Allred, B. 2020. When the protection of threatened species depends on the economy of foreign nations. PLoS One, 15(3): e0229555.
  24. Jones, P.F., Jakes, A.F., Eacker, D.R., Seward, B.C., Hebblewhite, M. & Martin, B.H. 2020. Evaluating Responses by Sympatric Ungulates to Fence Modifications across the Northern Great Plains. Wildlife Society Bulletin, Early View, Feb 2020,


  1. DeCesare, N., Weckworth, B.V., Walker, A., Bergman, E., Colson, K., Corrigan, R., Harris, R., Hebblewhite, M., Jesmer, B., Newby, J., Smith, J.R., Tether, R.M., Thomas, T., Pilgrim, K.L. & Schwartz, M. 2019. Phylogeography of a range edge subspecies: is there such thing as Shiras moose. Journal of Mammalogy, In Press.
  2. Squires, J.R., Olson, L., Heinemeyer, K., Hebblewhite, M. & Holbrook, J. 2019. Winter recreation and Canada lynx: reducing conflict through niche partitioning. Ecosphere, 10(10):e02876. 10.1002/ecs2.2876.
  3. Jones, P., Jakes, A., Sawyer, H., Telander, A., Martin, B., Hebblewhite, M. 2019. Fences reduce habitat for a partially migratory ungulate in the Northern Sagebrush Steppe. Ecosphere, 10(7):e02782. 10.1002/ecs2.2782.
  4. Muhly, T., Johnson, C.A., Hebblewhite, M., Neilson, E.W., Fortin, D., Fryxell, J.M., Latham, A.D., Latham, M.C., McLoughlin, P.D., Merrill, E.H., Paquet, P.C., Patterson, B.R., Schmiegelow, F., Scurrah, F., Musiani, M. 2019. Functional responses of wolves to human development across Boreal ecosystems. Ecology and Evolution, 1: 1-15,
  5. Hebblewhite, M. 2019. Wild Migrations: Atlas of Wyoming's Ungulates. Journal of Wildlife Management, 83(7):1628–1629.
  6. Berg, J.E., Hebblewhite, M., Cassady St. Clair, C. & Merrill, E.H. 2019. Prevalence and mechanisms of partial migration in ungulates. Frontiers in Ecology and Evolution, Front. Ecol. Evol. 7:325., doi: 10.3389/fevo.2019.00325
  7. Tempa, T., Mills, L.S., Hebblewhite, M. 2019. The spatial distribution and population density of Bengal tigers in Bhutan. Biological Conservation, 238: 108192,
  8. Heinemeyer, K., Squires, J., Hebblewhite, M., Smith, J., Holbrook, J.D. & Copeland, C. 2019. Wolverines in winter: Indirect habitat loss and functional responses to backcountry recreation. Ecosphere, 10, e02611.
  9. Steenweg, R., Hebblewhite, M., McKelvey, K., Lukacs, P. & Whittington, J. 2019. Species-specific trade-offs in statistical power when monitoring trends in multispecies occupancy. Ecosphere, 10, e02639.
  10. Cavedon, M., Goublin, C., Heppenheimer, E., von Holdt, B., Mariani, S., Hebblewhite, M., Hegel, T.M., Hervieux, D., Serrouya, R., Steenweg, R., Weckworth, B.V. & Musiani, M. 2019. Genes, environment and balancing selection in behaviorally bimodal populations: the caribou case. Molecular Ecology, 28, 1946-1963.
  11. Proffitt, K.M., DeVoe, J.D., Barker, K.J., Durham, R., Hayes, T., Hebblewhite, M., Jourdonnais, C., Ramsey, P. & Shamhart, J. 2019. A century of changing fire management alters ungulate forage in a wildfire-dominated landscape. Forest Ecology and Management, 44, 1-17.


  1. Eacker, D.R., Hebblewhite, M., Steenweg, R., Russell, M., Flasko, A. & Hervieux, D. (2018) Web-based application for estimating threatened woodland caribou population trends. Wildlife Society Bulletin, EarlyView Online.
  2. Buchanan, G. M., A. E. Beresford, M. Hebblewhite, F. J. Escobedo, H. M. de Klerk, P. F. Donald, P. Escribano, L. P. Koh, J. Martinez-Lopez, N. Pettorelli, A. K. Skidmore, Z. Szantoi, K. Tabor, M. Wegmann, and S. Wich. 2018. Free satellite data key to conservation. Science 361:139-140. DOI: 10.1126/science. aau2650
  3. Buchanan, G. M., A. E. Beresford, M. Hebblewhite, F. J. Escobedo, H. M. de Klerk, P. F. Donald, P. Escribano, L. P. Koh, J. Martinez-Lopez, N. Pettorelli, A. K. Skidmore, Z. Szantoi, K. Tabor, M. Wegmann, and S. Wich. 2018. Free satellite data key to conservation. Science 361:139-140. DOI: 10.1126/science. aau2650
  4. Weckworth, B., Hebblewhite, M., Mariani, S., Musiani, M. 2018. Lines on a map: conservation units, meta-population dynamics and recovery of woodland caribou in Canada. Ecosphere, 9(7): e02323.
  5. Finnegan, L., Pigeon, K., Cranston, J., Hebblewhite, M., Musiani, M., Neufeld, L., Schmiegelow, F., Duval, J. Stenhouse, G. 2018. Natural regeneration on seismic lines influences movement behavior of wolves and grizzly bears. PLoS One 13(4): e0195480. 
  6. Kosterman, M., Squires, J.R., Holbrook, J., Hebblewhite, M. 2018. Connectivity creates kittens: landscape pattern versus maternal state influences reproductive success of Canada lynx. Ecological Applications, 28(4): 1032 – 1043.
  7. Jakes, A., Gates, C.C., DeCesare, N.J., Jones, P., Goldberg, J.F ., Kunkel, K., Hebblewhite, M. 2018. Classifying the migration behaviors of pronghorn on their northern range. Journal of Wildlife Management. 82(6); 1229 – 1242. DOI: 10.1002/jwmg.21485. 
  8. Lukacs, P.M., Hebblewhite, M., Johnson, B.K., Johnson, H.E., Kauffman, M., Proffitt, K.M., Zager, P., Hersey, K., Holland, A.A., Hurley, M ., McCorquodale, S., Middleton, A., Mitchell, M.S., Nordhagen, M., Nowak, J., Walsh, J.J., White, P.J., 2017. Recovering large carnivores reduce juvenile elk recruitment at regional scales amid widespread declines. Journal of Wildlife Management, 82(4): 698 – 710. DOI: 10.1002/jwmg.21438.
  9.  Squires, J., Heinemeyer, K., Hebblewhite, M. 2018. A study of shared habitats: tracking forest carnivores and backcountry recreationists. The Wildlife Professional, 12.1, 45-49.
  10. Hebblewhite, M., Hurley, M ., Lukacs, P., Nowak, J. 2018. Predicting mule deer ( Odocoileus hemionus ) harvest in real-time: Integrating satellite remote-sensing measures of forage quality and climate in Idaho, United States. Pages 194-228 in Buchanan, G., and Leidner, A.K. (editors), Satellite Remote Sensing for Conservation Action: Case Studies of Implementation, Cambridge University Press.


  1. Steenweg, R., Hebblewhite, M., Whittington, J., McKelvey, K., Lukacs, P. 2017. Sampling scales define occupancy and the occupancy-abundance relationship in animals, Ecology, Earlyview Online: DOI: 10.1002/ecy.2054 
  2. Olson, L. E., J. Squires, E. Roberts, A. Miller, J. S. Ivan, and M. Hebblewhite. 2017. Modeling large-scale winter recreation terrain selection with implications for recreation management and wildlife. Applied Geography, 86: 66-91. 
  3. Jones, P., Jakes, A., Eacker, D., Seward, B., Hebblewhite, M., Martin, B. 2018. Evaluating responses by pronghorn to fence modifications across the Northern Great Plains. Wildlife Society Bulletin, April 10, 2018.
  4. Tucker, M.A., Bohning-Gaese, K., Fagan, W.F., Fryxell, J.M., Van Moorter, B., Alberts, S.C., Ali, A.H., Allen, A.M., Attias, N., Avgar, T., Bartlam-Brooks, H.L.A., Bayarbaatar, B., Belant, J.L., Bertassoni, A., Beyer, D.E., Bidner, L., van Beest, F.M., Blake, S., Blaum, N., Bracis, C., Brown, D., de Bruyn, P.N., Cagnacci, F., Calabrese, J.M., Camilo-Alves, C., Chamaille-Jammes, S., Chiaradia, A., Davidson, S.C., Dennis, T., DeStefano, S., Deifenbach, D., Douglas-Hamilton, I., Fennesey, J., Fichtel, C., Fiedler, W., Fischer, C., Fischoff, I., Fleming, C.H., Ford, A.T., Fritz, S., Gehr, B., Goheen, J.R., Gurarie, E., Hebblewhite, M., Heurich, M., Hewison, A.J., Hof, C., Hurme, E., Isbell, L.A., Janssen, R., Jeltsch, F., Kaczensky, P., Kane, A., Kappler, P., Kauffman, M., Kays, R., Kimuyu, D., Koch, F., Kranstauber, B., Lapoint, S.D., Leimgruber, P., Linnell, J.D.C., Lopez-Lopez, P., Markham, A.C., Mattison, J., Medici, E.P., Mellone, U., Merrill, E.H., de Miranda-Mourao, G., Morato, R.G., Morellet, N., Morrison, T.A., Diaz-Munoz, S.L., Mysterud, A., Nandintsetseg, D., Nathan, R., Niamir, A., Odden, J., O'Hara, R.B., Olvieria-Santos, G.R., Olson, K.A., Patterson, B.D., de Paula, R.C., Pedrotti, L., Rimmler, M., Rogers, T.L., Rolandsen, C.M., Rosenberry, C.S., Rubenstein, D.I., Safi, K., Said, S., Sapir, N., Sawyer, H., Schmidt, N.M., Selva, N., Sergiel, A., Shiilegdamba, E., Silva, J.P., Singh, N.J., Solberg, E.J., et al. 2017. Moving in the anthropocene: global reductions in terrestrial mammalian movements. Science, 359, 466-469.  10.1126/science.aam9712
  5. Whittington, J., Hebblewhite, M., Chandler, R. 2017. Generalized spatial mark-resight models for efficient density estimation of partially marked populations: an application to grizzly bears in the Rocky Mountains. Journal of Applied Ecology, early view. doi: 10.1111/1365-2664.12954 
  6. Olson, L. E., J. Squires, E. Roberts, A. Miller, J. S. Ivan, and M. Hebblewhite. 2017. Modeling large-scale winter recreation terrain selection with implications for recreation management and wildlife. Applied Geography, 86: 66-91. doi: 10.1016/j.apgeog.2017.06.023
  7. Hebblewhite, M. 2017. "North American Energy Sprawl and Triage Conservation: A Tale of Two Ecosystems," Ch. 3 in Kisecker, J., and Naugle, D. 2017. Energy Sprawl Solutions, Island Press. June 2017. 
  8. Jedrzejewska, W., Goldberg, J.F., Hebblewhite, M., Abarca, M., Gamarra, G., Calderon, L.E., Romero, J.F., Viloria, A.L., Carreno, R., Robinson, H.S., Lampo, M., Boede, E., Biganzoli, A., Stachowicz, Velasquez, G. 2017. Density and population structure of the jaguar (Panthera onca) in Hato Piñero, Los Llanos, Venezuela: A methodological comparison with long-term monitoring. Mammal Review62: 9-19.
  9. Eacker, D.R., Lukacs, P.M., Proffitt, K.M., & Hebblewhite, M. 2017. Assessing the importance of demographic parameters for population dynamics using Bayesian integrated population modeling. Ecological Applications. 10.1002/eap.1521


  1. Spitz, D., Hebblewhite, M., Stephenson, T.R. 2016. MigrateR: extending model-driven methods for classifying and quantifying animal movement behavior. Ecography 10.1111/ecog.02587
  2. Hurley, M., Hebblewhite,M., Lukacs, P.M., Nowak, J., Gaillard, J.-M., Bonenfant, C., 2016. Regional-scale models for predicting overwinter survival of juvenile ungulates. Journal of Wildlife Management. 81. no. 3: 364-378, 10.1002/jwmg.21211
  3. Peters, W., Hebblewhite, M., Mysterud, A., Spitz, D., Focardi, S., Urbano, F., Morellet, N., Huerich, M., Kjellander, P, Linnell, J.D.C., Cagnacci, F. 2017. Migration in geographic and ecological space by a large herbivore. Ecological Monographs, 10.1002/ecm.1250
  4. Gubili, C., S. Mariani, B. V. Weckworth, P. Galpern, A. D. McDevitt, M. Hebblewhite, B. Nickel, and M. Musiani. 2017. Environmental and anthropogenic drivers of connectivity patterns: A basis for prioritizing conservation efforts for threatened populations. Evolutionary Applications 10:199-211.
  5. Ossi, F., Gaillard, J.-M., Hebblewhite, M., Morellet, N., Ranc, N., Sandfort, R., Kroeschel, M.Kjellander, P., Mysterud, A., Linnell, J., Heurich, M., Sonnichsen, L., Sustr, P., Berger, A., Rocca, M., Urbano, F., Cagnacci, F. 2017. Plastic response by a small cervid to ad-libitum supplemental feeding in winter across a wide environmental gradient. Ecosphere, 8(1): e01629
  6. Hebblewhite, M. 2017. Billion dollar boreal woodland caribou and the biodiversity impacts of the global oil and gas industry. Biological Conservation 206:102-111.
  7. Ahrestani, F.S., Hebblewhite, M., Smith, B., Running, S.W., Post, E., 2016. Dynamic complexity and stability of herbivore populations at the species distribution scale. Ecology, 97(11): 3184-3194.
  8. Roffler, G., Adams, L.G., Hebblewhite M. 2016. Summer habitat selection by Dall’s sheep in Wrangell-St. Elias National Park and Preserve, Alaska. Journal of Mammalogy, 98(1): 94-105, 10.1093/jmammal/gyw135
  9. Steenweg, R., Hebblewhite, M., Kays, R., Ahumada, J., Fisher, J., Burton, A., Townsend, S., Carbone, C., Rowcliff, M., Whittinton, J., Brodie, J., Royle, J.A., Switalski, A., Clevenger, A., Heim, N., Rich, L., 2016. Scaling up camera traps - monitoring the planet's biodiversity with networks of remote sensors. Frontiers in Ecology and Environment, 15(1): 26-34. 10.1002/fee.1448
  10. Eacker, D.R., Hebblewhite, M, Proffitt, K.M., Jimenez, B.S, Mitchell, M.S., & Robinson, H.S. 2015. Annual elk calf survival in a multiple carnivore systemJournal of Wildlife Management. 80(8): 1345-1359, DOI: 10.1002/jwmg.21133 [Featured Article!]
  11. Hauer, R., Locke, H., Dreitz, V., Hebblewhite, M., Lowe, W., Muhlfeld, C., Nelson, C., Proctor, M., Rood, S. 2016. Gravel-bed river floodplains: the ecological nexus of glaciated mountain landscapes. Science Advances, 2:e1600026, 10.1126/sciadv.1600026
  12. Steenweg, R., Hebblewhite, M., Lukacs, P., Whittington, J., Forshner, A., Johnston, B., Shepherd, B., Peterson, D. 2016. Camera-based occupancy monitoring at large scales: power to detect trends in grizzly bears across the Canadian Rockies. Biological Conservation, 201: 192 – 200. 
  13. Xiao, W., Feng, L., Mou, P., Miquelle, D.G., Hebblewhite, M., Goldberg, J.F., Robinson, H.S., Zhao, X., Zhou, B., Wang, T., Ge, J. 2016.  Estimating abundance and density of Amur tigers along the Sino-Russian border. Integrative Zoology, 11(4): 322-332.
  14. Proffitt, K., Hebblewhite, M., Peters, W., Hupp, N., Shamhart, J. 2016. Linking landscape-scale differences in forage to ungulate nutritional ecology. Ecological Applications, 26(7), 2016, pp. 2156–2174.
  15. Eggeman, S., Hebblewhite, M., Bohm, H., Whittington, J., Merrill, E.H. 2016. Behavioral Flexibility in Migratory Behavior in a Long-lived UngulateJournal of Animal Ecology, 85: 785-797.
  16. Mancinelli, S., W. Peters, L. Boitani, F. Cagnacci. Roe deer summer habitat selection at multiple spatio-temporal scales in an Alpine environment, Hystrix (Italian Journal of Mammalogy) 26: 132-140.
  17. Steenweg, R., M. Hebblewhite, D. Gummer, B. Low, and B. Hunt. 2016. Assessing Potential Habitat and Carrying Capacity for Reintroduction of Plains Bison (Bison bison bison) in Banff National Park. PLoS One. DOI: 10.1371/journal.pone.0150065


  1. Guangshun J., Qi, J., Wang, G. , Shi, Q., Darman, Y., Hebblewhite, M., Miquelle, D., Li, Z., Zhang, X., Gu, J., Chang, Y., Zhang, M., Ma, J.. 2015. New hope for the survival of the Amur leopard in China. Scientific Reports, 5:15475,  DOI: 10.1038/srep15475
  2. Proffitt, K. M., J. F. Goldberg, M. Hebblewhite, R. Russell, B. Jimenez, H. Robinson, K. Pilgrim, and M. K. Schwartz. 2015. Integrating resource selection and harvest into spatial capture-recapture models for large carnivoresEcosphere. DOI: 10.1890/ES15-00001.1 
  3. Goldberg, J.F., Tempa, T., Norbu, N., Hebblewhite, M., Mills, L.S., Wangchuk, T. 2015. First estimates of common leopard density in the lower foothills of Bhutan. PLoS One, DOI: 10.1371/journal.pone.0140757
  4. Cagnacci, F., S. Focardi, A. Ghisla, B. van Moorter, E. Gurarie, M. Heurich, R. May, T. Nygard, A. Mysterud, J. Linnell, M. Panzacchi, E. H. Merrill, C. M. Rolandsen, and M. Hebblewhite. 2015. How many routes lead to migration? Re-establishing the link between definitions, methods and movement ecologyJournal of Animal Ecology, 85(1): 54-68. doi: 10.1111/1365-2656.12449
  5. DeCesare, N.J., Hebblewhite, M., Lukacs, P., Hervieux, D. 2015. Evaluating common sources of censoring and truncation in telemetry-based survival data. Journal of Wildlife Management. 80(1): 138-148. DOI: 10.1002/jwmg.991
  6. Robinson, H., Ruth, T.K., Gude, J., Choate, D., DeSimone, R., Hebblewhite, M., Kunkel, K., Matchett, R., and Murphy, K. 2015. Linking Resource Selection and Mortality Modeling for Population Estimation of Mountain Lions in MontanaEcological Modeling, 312: 11-25.
  7. Latham, A.D., Latham, M.C., Anderson, D.P., Cruz, J., Herries, D., Hebblewhite, M. 2015. The GPS craze: six questions to address before deciding to deploy GPS technology on wildlife. New Zealand Journal of Ecology. 39(1): 0-0.


  1. Ossi, F., J. M. Gaillard, M. Hebblewhite, and F. Cagnacci. 2014. Snow sinking depth and forest canopy drive winter resource selection more than supplemental feeding in an alpine population of Roe deer. European Journal of Wildlife Research, 61: 111-124.
  2. Morgan-Henderson, M., M. Hebblewhite, J. Stetz, K. Kendall, and R. Carlson. 2015. Modeling multi-scale resource selection for bear rubs in northwestern Montana and Idaho. Ursus. 26(1): 28-39. doi: 10.2192/URSUS-D-14-00026.1
  3. Peters, W., Cavedon, M., Hebblewhite, M., Pedrotti, L., Mustoni, A., Zibordi, F., Groff, C., Zanin, M., Cagnacci, F., 2015. Resource selection and connectivity reveal conservation challenges for reintroduced brown bears in the Italian Alps. Biological Conservation, 186: 123-133.
  4. Berger, J., G. B. Schaller, E. Cheng, A. Kang, M. Krebs, L. Li, and M. Hebblewhite. 2015. Legacies of past exploitation and climate affect mammalian sexes differently on the roof of the world. Scientific Reports, 5 : 8676 | DOI: 10.1038/srep08676
  5. Hervieux, D., M. Hebblewhite, N. J. DeCesare, S. Boutin, M. Russell, and S. Robertson. 2015. Reply to the comment by Harron on"Widespread and drastic declines in woodland caribou (Rangifer tarandus cairbou) continue unabated in Alberta". Canadian Journal of Zoology, 93: 1–4. (open access)
  6. Hervieux, D., M. Hebblewhite, M. M. Bacon, Stepnisky, D., and S. Boutin. 2014. Managing wolves to recover threatened woodland caribou in Alberta. Canadian Journal of Zoology, 92(12): 1029-1037, 10.1139/cjz-2014-0142
  8. Semeniuk, C.A.D., Musiani, M., Birkigt, D., Hebblewhite, M., Grindal, S., and Marceau, D.J. 2014. Identifying non-independent anthropogenic risks using a behavioral individual-based modelEcological Complexity,17 (2014) 67–78
  9. Brodie, J., Giordano, A., Dickson, B.G., Hebblewhite, M., Bernard, H., Mohd-Azlan, J., Anderson, J., and Ambu, L. 2014. Evaluating multispecies landscape connectivity in a threatened tropical community. Conservation Biology, EarlyView Online, DOI: 10.1111/cobi.12337
  10. Whittington, J., K. Heuer, B. Hunt, and M. Hebblewhite. 2014. Multi-species occupancy using spatially and temporally replicated snow surveys. Animal Conservation, EarlyView Online, doi:10.1111/acv.12140
  11. Robinson, H., R. DeSimone, C. Hartway, J. A. Gude, M. Thompson, M. S. Mitchell, and M. Hebblewhite. 2014. A test of the compensatory mortality hypothesis in mountain lions: a management experiment in west-central Montana. Journal of Wildlife Management, 78: 791-807. DOI: 10.1002/jwmg.726
  12. Goldberg, J.F., Hebblewhite, M., Bardsley, J. 2014. Consequences of a Refuge for the Predator-Prey Dynamics of a Wolf-Elk system in Banff National Park, Alberta, Canada. PLoS One 9(3): e91417.
  13. Hurley, M., Hebblewhite, M., Gaillard, J.M., Dray, S., Taylor, K., Smith, B., Zager, P., and Bonenfant, C. 2014. Functional analysis of NDVI curves reveals overwinter mule deer survival is driven by both spring and fall phenology, not just spring. Philosophical Transactions of the Royal Society B: Biological Sciences, 369: 20130196.
    1. Electronic Supplementary Materials (including R code, data in the paper, and background on FPCA methods)
  14. Hebblewhite, M., D. G. Miquelle, H. Robinson, A. A. Murzin, D. G. Pikunov, and Y. M. Dunishenko. 2014. Including biotic interactions with ungulate prey and humans improves habitat conservation modeling for endangered Amur Tigers in the Russian Far East. Biological Conservation, 178: 50-64.
  15. Peters, W., Hebblewhite, M., Smith, K. G., Webb, S. M., Webb, N. & Anderson, R. 2014. Moose Population Estimation Using Distance Sampling in West-Central Alberta. Wildlife Society Bulletin, 38: 639-649.
  16. Ralmer, J., Kellenberg, D., Hebblewhite, M., Sime, C. 2014. Crying wolf: a spatial analysis of wolf locations and depredations on calf weight. American Journal of Agricultural Economics, 1–26; doi: 10.1093/ajae/aat100
  17. Ripple, W.J., Estes, J.A., Beschta, R.L., Wilmers, C.C., Ritchie, E.G., Hebblewhite, M., Berger, J., Elmhagen, B., Letnik, M., Nelson, M.P., Schmitz, O.J., Smith, D.W., Wallach, A.D., and Wirsing, A.J. 2014. Status and Ecological effects of the world's largest carnivores. Science 343: 123148. doi:DOI: 10.1126/science.1241484.


  1. Ahrestani, F.S.Hebblewhite, M., and Post, E.S. 2013. The importance of observation versus process error in analyses of global ungulate populations. Scientific Reports, 3: 03125, 3, doi:10.1038/srep03125.
  2. Hervieux, D., M. Hebblewhite, N. J. DeCesare, S. Boutin, M. Russell, and S. Robertson. 2013.Widespread and drastic declines in woodland caribou (Rangifer tarandus cairbou) continue unabated in Alberta. Canadian Journal of Zoology, 91: 872–882. (open access)
  3. DeCesare, N. J., M. Hebblewhite, M. Bradley, D. Hervieux, and M. Musiani. 2013. Linking habitat selection and predation risk to spatial variation in fitness. Journal of Animal Ecology Online early (open access). 
  4. Weckworth, B. V., M. Musiani, N. J. DeCesare, A. McDevitt, M. Hebblewhite, and S. Mariani. 2013. Preferred habitat and effective population size drive the landscape genetics of an endangered ungulate. Proceedings of the Royal Society B-Biological Sciences 280, 20131756
  5. Post, E. S., E. Bhatt, C. Bitz, J. Brodie, T. L. Fulton, M. Hebblewhite, J. Kerby, S. Kutz, I. Stirling, and D. A. Walker. 2013. Sea ice as a driver of ecological responses to climate change in the Arctic. Science 341 no. 6145 pp. 519-524
  6. Polfus, J. L., K. Heinemeyer, M. Hebblewhite, and Taku River Tlingit First Nation. 2013. Comparing Traditional Ecological Knowledge and Western Science Woodland Caribou Habitat ModelsJournal of Wildlife Management, 78(1):112-121. doi: 10.1002/jwmg.643
  7. Hebblewhite, M. 2013. Consequences of ratio-dependent wolf predation on elk population dynamics. Population Ecology, 55(4): 511-522.
  8. Latham, A., Latham, M., Knopff, K., Hebblewhite, M., Boutin, S. 2013. Wolves, white-tailed deer, and beaver: implications of seasonal prey switching for woodland caribou declines. Ecography,36(12): 1276-1290. doi: 10.1111/j.1600-0587.2013.00035.x
  9. Muhly, T.B., Hebblewhite, M., Paton, D., Pitt, J.A., Boyce, M.S., Musiani, M. 2013. Pervasive human influences in a terrestrial food web put to rest the top-down versus bottom-up debate. PLoS One, 8:e64311.
  10. Miller, C. S., M. Hebblewhite, Y. Petrunenko, I. V. Seryodkin, N. J. DeCesare, J. M. Goodrich, and D. G. Miquelle. 2013. Estimating Amur tiger (Panthera tigris altaica) kill rates and consumption rates using GPS collars. Journal of Mammalogy, 94:000-000.
  11. Brodie, J., H. E. Johnson, M. S. Mitchell, P. Zager, K. M. Proffitt, M. Hebblewhite, M. Kauffman, B. Johnson, J. Bissonette, C. Bishop, J. A. Gude, J. Herbert, K. Hersey, M. Hurley, P. Lukacs, S. McCorquodale, E. McIntire, J. Nowak, D. W. Smith, and P. J. White. 2013. Relative influence of human harvest, carnivores, and weather on adult female elk survival across western North America Journal of Applied Ecology, 50(2): 295-305. doi: 10.1111/1365-2664.12044
  12. Tempa, T.M. Hebblewhite, L. S. Mills, N. Norbu, T. Wangchuk, T. Wangchuk, T. Nidup, P. Dendup, D. Wangchuk, Y. Wangdi, and T. Dorji. 2013. Royal Manas National Park, Bhutan, as a wild felid biodiversity hot spotOryx, 47(2): 207-210.  
  13. Johnson, H. E., M. Hebblewhite, T. R. Stephenson, D. W. German, and B. M. Pierce. 2013.Apparent competition limits the recovery of an endangered ungulate: quantifying the direct and indirect effects of predation. Oecologia 171:295–307.


  1. Greene, L., M. Hebblewhite, and T. R. Stephenson. 2012. Short-term vegetation response to wildfire in the eastern Sierra Nevada: implications for recovering an endangered ungulate. Journal of Arid Environments, 87: 118 – 128
  2. Peters, W., Hebblewhite, M., DeCesare, N.J., Cagnacci, F., Musiani, M., 2012. Human landscape alteration affects resource partitioning of coexisting moose and threatened caribouEcography, 35: 001-012.
  3. Cleveland, S. M., M. Hebblewhite, M. Thompson, and R. Henderson. 2012. Linking elk movement and resource selection to hunting pressure in heterogeneous landscapes. Wildlife Society Bulletin, 36:658–668.
  4. Squires, J. R., N. J. DeCesare, L. E. Olson, J. A. Kolbe, M. Hebblewhite, and S. Parks. 2012.Combining resource selection and movement behavior to predict corridors for Canada lynx at their southern range periphery. Biological Conservation, 157: 187 - 195.
  5. Squires, J.R., DeCesare, N.J., Hebblewhite, M., Berger, J. 2012. Missing Lynx and Trophic Cascades in Food Webs: a Reply to Ripple et al. Wildlife Society Bulletin, 36: 567 – 571.
  6. Semeniuk, C. A. D., M. Musiani, M. Hebblewhite, S. Grindal, and D. J. Marceau. 2012.Incorporating behavioral-ecological strategies in pattern-oriented modeling of caribou habitat use in a highly industrialized landscape. Ecological Modeling 243: 18 – 32.
  7. Hebblewhite, M., F. Zimmermann, Z. Li, D. G. Miquelle, Z. Minghai, S. Haiyi, F. Morschel, W. Zhigang, S. Lianxi, A. Purekhovsky, and Z. Chunquan. 2012. A Future for Amur Tigers in Northeastern China? Predicting Potential Tiger Habitat and Numbers in the Changbaishan EcosystemAnimal Conservation, 15 :579–592.
  8. Weckworth, B. V., M. Musiani, A. McDevitt, M. Hebblewhite, and S. Mariani. 2012. Reconstruction of caribou evolutionary history in Western North America and its implications for conservation. Molecular Ecology 21: 3610-3624.
  9. Kohl, M., S. M. ClevelandM. Hebblewhite, and R. C. Callaway. 2012. Forage Value of Invasive Species to the Diet of Rocky Mountain Elk. Rangelands 34: 24-28.
  10. DeCesare, N. J., M. Hebblewhite, F. Schmiegelow, D. Hervieux, G. J. McDermid, L. Neufeld, M. Bradley, J. Whittington, K. G. Smith, L. E. Morgantini, M. Wheatley, and M. Musiani. 2012.Transcending scale dependence in identifying habitat with resource selection functions. Ecological Applications 22: 1068-1083.
  11. Nelson, B.Hebblewhite, M., Ezenwa, V., Shury, T., Merrill, E. H., Paquet, P. C., Schmiegelow, F., Seip, D., Skinner, G. & Webb, N. 2012. Seroprevalence of canine parvovirus and distemper in wolves in the Canadian Rocky MountainsJournal of Wildlife Diseases, 48: 68-78.
  12. Mitchell, M. S. and Hebblewhite, M. 2012. Carnivore habitat ecology: integrating theory and application for conservation. Pages 218 - 255 in R. A. Powell and L. Boitani, editors. Carnivore Ecology and Conservation: A Handbook of Techniques. Oxford University Press, Oxford, UK.
  13. Boutin, S., M. S. Boyce, M. Hebblewhite, D. Hervieux, K. H. Knopff, M. C. Latham, D. M. Latham, J. Nagy, D. Seip, and R. Serrouya. 2012. Why are caribou declining in the oil sands? Frontiers in Ecology and Environment 10:65-67.
  14. Robinson, H. S.Hebblewhite, M., DeCesare, N. J., Whittington, J., Neufeld, L., Bradley, M. & Musiani, M. 2012. The effect of fire on spatial separation between wolves and caribou. Rangifer: 20: 277–294.
  15. DeCesare, N.J., Hebblewhite, M., Bradley, M., Smith, K., Hervieux, D. & Neufeld, L. 2012.Estimating ungulate recruitment and growth rates using age ratiosJournal of Wildlife Management 76: 144-153.


  1. Miller, C. S., Petrunenko, Y. K., Goodrich, J. M., Hebblewhite, M., Seryodkin, I. V. & Miquelle, D. G. 2011 Translocation a success, but poaching remains a problem for Amur tigers. Cat News, 55: 23-26.
  2. Rogala, J. K., M. Hebblewhite, J. Whittington, C. A. White, J. Coleshill, and M. Musiani. 2011.Human Activity Differentially Redistributes Large Mammals in the Canadian Rockies National Parks. Ecology and Society, 16(3): 16.
  3. Hebblewhite, M. & Merrill, E. H. 2011. Fitness balancing in a partially migratory ungulate through risk-forage trade-offs. Oikos, 120(12): 1860 - 1870.
  4. Polfus, J.L.Hebblewhite, M., Heinemeyer, K. 2011. Identifying indirect habitat loss and avoidance of human infrastructure by northern mountain caribou. Biological Conservation, 144:2637-2646. Supplementary materials.
  5. Hebblewhite, M. 2011. Unreliable Knowledge about Economic Impacts of Large Carnivores on Bovine Calves. Journal of Wildlife Management, 75(8): 1724-1730. doi: 10.1002/jwmg.206
  6. Whittington, J., Hebblewhite, M., DeCesare, N. J., Neufeld, L., Bradley, M., Wilmshurst, J. W. & Musiani, M. 2011. Caribou encounters with wolves increases near roads and trails: a time-to-event approach. Journal of Applied Ecology, 48: 1535-1542. doi: 10.1111/j.1365-2664.2011.02043.x
  7. Hebblewhite, M., Miquelle, D. G., Aramilev, V. V. & Pikonov, D. G. 2011. Recovering the critically endangered Far Eastern Amur Leopard: identifying potential habitat and population size for reintroduction. Biological Conservation, 144:2403-2413. Supplementary materials
  8. Hurley, M.Hebblewhite, M., Unsworth, J., Zager, P., Garton, E., Montgomery, D., Skalski, J. and C. Maycock. 2011. Mule deer demographic response of mule deer to experimental reduction of coyotes and mountain lions in southeastern Idaho. Wildlife Monographs, 178: 1 -33.
  9. Vucetich, J. A., Hebblewhite, M., Smith, D. W. & Peterson, R. O. 2011. Predicting prey population dynamics from kill rate, predation rate and predator-prey ratios in three wolf-ungulate systems. Journal of Animal Ecology, 80: 1236 - 1245. Supplementary materials
  10. Griffin, K., Hebblewhite, M., Zager, P., Robinson, H. R., Barber-Meyer, S., Christenson, D., Creel, S., Harris, N., Hurley, M., Jackson, D., Johnson, B., Mech, L. D., Myers, W. L., Raithel, J., Schlegel, M., Smith, B. & White, P. J. 2011. Neonatal mortality of elk driven by climate, predator phenology and predator diversity. Journal of Animal Ecology, 80: 1246 - 1257. Supplementary materials.
  11. DeCesare, N.J., Whittington, J., Robinson, H., Hebblewhite, M., Bradley, M., Neufeld, L. & Musiani, M. 2011. The role of translocation in recovery of woodland caribou populations. Conservation Biology, 25: 365 - 373.
  12. Miller, C. S., Hebblewhite, M., Goodrich, J. M. & Miquelle, D. G. 2010 Review of research methodologies for tigers - Telemetry. Integrative Zoology, 5: 378 - 389.
  13. Hebblewhite, M. 2011. Effects of Energy Development on Ungulates in Western North America. Pages 71-94 in D. Naugle, editor. Energy Development & Wildlife Conservation in Western North America. Island Press, New York.


  1. Matthiopoulos, J., Hebblewhite, M., Aarts, G. & Fieberg, J. 2010. Generalised functional responses for species distributions. Ecology, 92: 583 - 589. Supplementary materials.
  2. Hebblewhite, M., Musiani, M. & Mills, L. S. 2010 Restoration of genetic connectivity among Northern Rockies wolf populations. Molecular Ecology, 19: 4383-4385.
  3. Musiani, M., Anwar, M. S., McDermid, G., Hebblewhite, M. & Marceau, D. J. 2010 How humans shape wolf behavior in Banff and Kootenay National Parks, Canada. Ecological Modeling 221: 2374 – 2387.
  4. Muhly, T. B., Alexander, M., Boyce, M.S., Creasey, R., Hebblewhite, M., Paton, M., Pitt, J. and M. Musiani. 2010. Differential risk effects of wolves on elk and cattle: implications for conservation. Oikos 119, 1243-1254.
  5. Gaillard, J. M., Hebblewhite, M., Loison,A., Fuller, F., and R. Powell. 2010. Habitat-performance relationships: finding the right metric at a given spatial scale. Philosophical Transactions of the Royal Society B-Biological Sciences 365: 2255-2265.
  6. Beyer H.L., Haydon D.T., Morales J.M., Frair J.L., Hebblewhite M., Mitchell M.S. & Matthiopoulos J. 2010. Habitat preference: understanding use versus availability designs. Philosophical Transactions of the Royal Society B-Biological Sciences, 365: 2245-2254.
  7. Merrill, E. H., Sand, H., Zimmerman, B., McPhee, H., Hebblewhite, M., Webb, N., Wabakkan, P. & Frair, J. L. 2010. Opportunities and challenges for using predator movements to assess kill sites and attack rates. Philosophical Transactions of the Royal Society B-Biological Sciences, 365:2279-2288.
  8. Frair, J. L., Fieberg, J., Cagnacci, F., Hebblewhite, M., Pedrotti, L. & DeCesare, N. 2010. Resolving issues of imprecise and habitat-biased locations in ecological analyses using GPS telemetry data
  9.  Philosophical Transactions of the Royal Society B-Biological Sciences, 365(1550):2187-2200. doi: 10.1098/rstb.2010.0084
  10. Fieberg, J., Matthiopoulos, J., Hebblewhite, M., Boyce, M. S. & Frair, J. L. 2010. Autocorrelation and studies of habitat selection: problem, red herring, or opportunity? Philosophical Transactions of the Royal Society B-Biological Sciences, 365: 2233-2244. 
  11. Hebblewhite, M. & Haydon, D. T. 2010. Distinguishing technology from ecology: a critical review of the use and abuse of GPS collar data in ecology. Philosophical Transactions of the Royal Society B-Biological Sciences, 365: 2303-2312.
  12. Robinson, B. G., M. Hebblewhite, and E. Merrill. 2010. Are migrant and resident elk (Cervus elaphus) exposed to similar forage and predation risk on their sympatric winter range? Oecologia 164: 265-275.
  13. DeCesare, N.J.Hebblewhite, M., Robinson, H., Musiani, M. 2010. Endangered, apparently: the role of apparent competition in endangered species management. Animal Conservation, 13: 353-362.


  1. Hebblewhite, M., White, C.A., & Musiani. M. 2009. Revisiting extinction in National Parks: Mountain caribou in Banff. Conservation Biology, 24: 341-344.
  2. Hebblewhite, M., & Merrill, E.H. 2009. Trade-offs between wolf predation risk and forage at multiple spatial scales in a partially migratory ungulate. Ecology, 90(12): 3445-3454.
  3. Hebblewhite, M. 2009. Linking wildlife populations with ecosystem change: state of the art satellite ecology for national park science. Park Science, 26(1): 54-59
  4. Eggeman, S., Hebblewhite, M., Cunningham, J., & Hamlin, K. 2009. Fluctuating asymmetry in elk antlers is unrelated to environmental conditions in the Greater Yellowstone Ecosystem. Wildlife Biology 15: 299-309.
  5. Post, E., Brodie, J., Wilmers, C.C., Hebblewhite, M., & Anders, A.D. 2009. Global population dynamics and hotpots of climate change. Bioscience, 59: 489-499.
  6. McDevitt, A., Mariani, S.L., Hebblewhite, M., DeCesare, N.J., L., Seip, D., Weckworth, B. V. & Musiani, M. 2009. Survival in the Rockies of an endangered hybrid swarm from diverged caribou (Rangifer tarandus) lineages. Molecular Ecology, 18: 665-679.
  7. Hebblewhite, M., Munro, R. & Merrill, E. H. 2009. Trophic consequences of post-fire logging in a wolf-elk system. Forest Ecology and Management, 257, 1053-1062.


  1. Hebblewhite, M., Merrill, E. H. & McDermid, G. 2008. A mutli-scale test of the forage maturation hypothesis for a partially migratory montane elk population. Ecological Monographs, 78: 141-166.
  2. Webb, N., Hebblewhite, M., Merrill, E.H. 2008. Identifying wolf kill sites using global positioning systems. Journal of Wildlife Management, 72: 798- 807.
  3. Hebblewhite, M., & Merrill, E.H. 2008. Modelling wildlife-human relationships for social species with mixed-effects resource selection models. Journal of Applied Ecology, 45: 834-844.Supplementary materials.


  1. Hebblewhite, M., Whittington, J., Bradley, M., Skinner, G., Dibb, A. & White, C. A. 2007.Conditions for caribou persistence in the wolf-elk-caribou systems of the Canadian Rockies. Rangifer, 17, 79-91.
  2. Hebblewhite, M. 2007. Predator-prey management in the National park context: lessons from a transboundary wolf-elk, moose, and caribou system. Transactions of the North American Wildlife Conference, 72, 348-365.
  3. Hebblewhite, M., Percy, M. & Merrill, E. H. 2007. Are all GPS collars created equal? Correcting habitat-induced bias using three brands in the Central Canadian Rockies. Journal of Wildlife Management, 71: 2026-2033.
  4. Hebblewhite, M. & Merrill, E. H. 2007. Multiscale wolf predation risk for elk: does migration reduce risk? Oecologia, 152, 377-387.


  1. Hurford, A., Hebblewhite, M., & Lewis, M.A. 2006. A spatially explicit model for the Allee effect: why wolves recolonize so slowly in Greater Yellowstone. Theoretical Population Biology. 70: 244-254
  2. Gillies, C.S., Hebblewhite, M., Nielsen, S.E., Krawchuk, M.A., Aldridge, C.L., Frair, J.L., Saher, D.J., Stevens, & C.E., Jerde. 2006. Application of random effects to the study of resource selection by animals. Journal of Animal Ecology. 75: 887-898.
  3. Hebblewhite, M., Merrill, E. H., Morgantini, L. E., White, C. A., Allen, J. R., Bruns, E., Thurston, L. & Hurd, T. E. 2006. Is the migratory behavior of montane elk herds in peril? The case of Alberta’s Ya Ha Tinda elk herd. Wildlife Society Bulletin, 34: 1280-1295


  1. Hebblewhite, M., Merril, E.H., & McDonald, T.E. 2005. Spatial decomposition of predation risk using resource selection functions: an example in a wolf-elk predator-prey system. Oikos, 111:101-111.
  2. Hebblewhite, M., White, C. A., Nietvelt, C., McKenzie, J. M., Hurd, T. E., Fryxell, J. M., Bayley, S. & Paquet, P. C. 2005 Human activity mediates a trophic cascade caused by wolves. Ecology, 86: 2135-2144.
  3. Hebblewhite, M. 2005. Predation by wolves interacts with the North Pacific Oscillation (NPO) on a western North American elk population. Journal of Animal Ecology, 74: 226-233.


  1. Hebblewhite, M., Paquet, P.C., Pletscher, D.H., Lessard, R.J. & Callaghan, C. 2004.Development and application of a ratio-estimator to estimate wolf-killing rates and variance in a multi-prey ecosystem. Wildlife Society Bulletin, 31:933-945.


  1. Hebblewhite, M., Percy, M., & Serrouya, R. 2003. Black bear survival and demography in the Bow Valley of Banff National Park. Biological Conservation, 112: 415-425.
  2. Hebblewhite, M., Pletscher, D.H., & P.C. Paquet. 2003. Elk population dynamics following wolf recolonization of the Bow Valley of Banff National Park. Parks Canada Research Links 11: 10-12.


  1. Hebblewhite, M., & Pletscher, D.H. 2002. Effects of elk herding on predation by wolves: linking anti-predator behavior to population dynamics. Canadian Journal of Zoology 80: 800-809.
  2. Hebblewhite, M., Pletscher, D.H., & P.C. Paquet. 2002. Factors affecting elk population growth rate in areas with and without predation by recolonizing wolves in Banff National Park. Canadian Journal of Zoology 80:789-799.


  1. Duke, D., Hebblewhite, M., Paquet, P.C., & Callaghan, C. 2001. Restoration of a large carnivore corridor in Banff National Park, Alberta. In Large mammal restoration: ecological and sociological challenges in the 21st Century, Edited by Maehr, D.S., Noss, R.F., & Larkin, J.L., Island Press.